Habit and leaf form. Herbs. Leaves well developed (with prominent green and white stripes, in non-flowering plants only, reduced to a minute bract in flowering plants), or absent (by misinterpretation). Plants succulent, or non-succulent. Perennial. Leaves basal. Tuberous. Mesophytic. Leaves small to medium-sized; ‘herbaceous’, or leathery, or fleshy; petiolate; simple. Leaf blades entire; flat; ovate; parallel-veined; cross-venulate, or without cross-venules. Leaf blade margins entire. Vernation conduplicate, or convolute. Leaf anatomy. Hairs absent (leaf smooth, shiny). Stem anatomy. Secondary thickening absent.
Reproductive type, pollination. Fertile flowers hermaphrodite. Unisexual flowers absent. Plants hermaphrodite. Entomophilous. Pollination mechanism conspicuously specialized.
Inflorescence and flower features. Flowers solitary; small to large; fragrant; very irregular; zygomorphic; resupinate. The floral asymmetry involving the perianth and involving the androecium. Flowers 3 merous; cyclic; supposedly basically pentacyclic. Perigone tube absent. Perianth of ‘tepals’, or with distinct calyx and corolla; 6; 2 -whorled; isomerous (but zygomorphic); free; spotted (labellum only); white to yellow (perianth members), or purple, green, and yellow (labellum). Calyx (if the outer whorl be so designated) 3 (the median member ostensibly posterior); 1 -whorled; polysepalous. Corolla (i.e. the members of the inner whorl) 3; polypetalous; imbricate. Androecium 3, or 1 (by misinterpretation). Androecial members free of the perianth; united with the gynoecium (fused with the style to form a column or ‘gynostemium’; column with red bands); coherent (via the gynostemium); 1 - adelphous; theoretically 2 -whorled. Androecium including staminodes, or exclusively of fertile stamens (by misinterpretation). Staminodes 2 (these anterior (ostensibly posterior), supposedly the abaxial pair of the inner whorl). Stamens 1 (this across the flower from the labellum, i.e. anterior but ostensibly posterior, supposedly representing the outer whorl); reduced in number relative to the adjacent perianth; alterniperianthial (i.e. with reference to the single stamen, across the flower from the labellum); filantherous, or with sessile anthers. Anthers dorsifixed to basifixed; dehiscing via longitudinal slits; introrse; tetrasporangiate; appendaged, or unappendaged. Pollen shed in aggregates; in the form of pollinia. Gynoecium 3 carpelled. The pistil 1 celled. Carpels isomerous with the perianth. Gynoecium syncarpous; eu-syncarpous; inferior. Ovary unilocular; 1 locular. The ‘odd’ carpel anterior (away from the labellum). Gynoecium stylate. Styles 1 (inflexed); apical. Stigmas 1; 3 - lobed (but becoming much modified in form, the apex of the median lobe forming the ‘rostellum’); wet type; papillate; Group III type. Placentation parietal. Ovules not differentiated; in the single cavity 30–100 (i.e. very numerous); non-arillate; anatropous.
Fruit and seed features. Fruit non-fleshy; dehiscent; a capsule. Capsules septicidal, or loculicidal. Fruit 30–500 seeded (i.e. seeds usually very numerous). Seeds endospermic (endosperm development arrested very early), or non-endospermic; minute; without starch. Embryo rudimentary at the time of seed release, or weakly differentiated. Seedling. Seedling collar not conspicuous. Primary root ephemeral.
Geography, cytology, number of species. Native of Australia. Endemic to Australia. Australian states and territories: Western Australia. South-West Botanical Province.
Additional characters Perianth of 5 similar members and the median inner member modified into the labellum (sepals and lateral petals narrowly ovate, tapering at the base; adaxial sepal erect, incurved; other segments spreading; labellum clawed, 3-lobed, c. half as long as the sepals, not fringed; mid-lobe longer than lateral lobes, with recurved apex, margin undulate). Leaves solitary. Column prominently winged. Labellum not insect-like. Perianth not glossy. Labellum with calli (lamina calli few to numerous, clubbed, in 2 parallel rows near apex).