Habit and leaf form. Trees, or shrubs; evergreen, or deciduous; laticiferous, or non-laticiferous and without coloured juice; resinous. Leaves cauline. Stem internodes solid. Self supporting, or climbing. Leaves alternate; spiral; ‘herbaceous’; petiolate (petiole more or less laterally ridged); flowers fragrant; simple. Leaf blades oblong (oblong-ovate), or obovate (to narrowly obovate or to oblong-obovate); veins prominent; cuneate at the base (narrowly, or decurrent). Leaves without stipules. Leaf blade margins entire. Leaf anatomy. Hairs present. Extra-floral nectaries absent. Stem anatomy. Secondary thickening developing from a conventional cambial ring.
Reproductive type, pollination. Fertile flowers hermaphrodite. Unisexual flowers absent. Plants hermaphrodite.
Inflorescence and flower features. Flowers aggregated in ‘inflorescences’; in panicles (usually). The terminal inflorescence unit racemose. Inflorescences terminal, or axillary. Flowers pedicellate (articulate just below the base of the calyx); bracteate (overlapping the base of the calyx); bracteolate (2 per flower); regular; 5 merous; cyclic; tetracyclic, or pentacyclic. Free hypanthium present (short), or absent. Hypogynous disk present; intrastaminal; annular. Perianth with distinct calyx and corolla; 3–5, or 6–10; 1 -whorled, or 2 -whorled; isomerous. Calyx present; (4–)5(–6); 1 -whorled; polysepalous, or gamosepalous; toothed (or ciliate); imbricate; regular. Sepals ovate (broadly; Note: this shape describes the free lobe of the joined calyx). Corolla present; (4–)5(–6); 1 -whorled; polypetalous; imbricate; white, or cream. Petals elliptic, or oblong (oblong-elliptic). Androecial members definite in number. Androecium 6. Androecial members free of the perianth; all equal; free of one another; 1 -whorled, or 2 -whorled. Androecium exclusively of fertile stamens, or including staminodes. Staminodes when present, 1–9. Stamens (8–)10(–12) (twice the number of petals); all more or less similar in shape; round the disc. Anthers basifixed; versatile; dehiscing via longitudinal slits; introrse; tetrasporangiate. Pollen shed as single grains. Gynoecium 4–5 carpelled. The pistil 1 celled. Gynoecium syncarpous (connate only at the base); synovarious; superior (usually). Ovary unilocular; 1 locular. Gynoecium stylate (the single fertile style is reduced). Styles 1 (1 fertile, 3–5 sterile); free; ‘gynobasic’ (having 3–5 sterile carpels whose the prominent stigmas function as gynobasic styles). Stigmas 5 (prominent and sterile, and 1 obsolete and fertile stigma). Placentation parietal, or basal. Ovules in the single cavity 1; pendulous, or ascending; apotropous; with ventral raphe, or with dorsal raphe; non-arillate; anatropous.
Fruit and seed features. Fruit fleshy (scarcely); indehiscent; a drupe; 1 seeded (per cell usually). Seeds non-endospermic. Cotyledons 2. Embryo curved.
Etymology. After Francis Hamilton (n?e Buchanan, 1762–1829).
Wheeler, J. R.; Rye, B. L.; Koch, B. L.; Wilson, A. J. G.; Western Australian Herbarium (1992). Flora of the Kimberley region. Western Australian Herbarium. Como, W.A.
Australia. Bureau of Flora and Fauna (1985). Flora of Australia. Volume 25, Melianthaceae to Simaroubaceae. Australian Govt. Pub. Service. Canberra.
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